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Despite some initial rivalries with them, she later became a trusted and knowledgeable member of Jack's West Jr's team. Iolanthe Compton-Jones was born as a royal. She and her older brother, Orlando , became members of the Kingdom of Land that included their British homeland, as part of the ancient group known as the Four Legendary Kingdoms.
Because of her membership in the Four Kingdoms, Iolanthe was taught about the true history of mankind, as well as learning of the trials set by super-ancient beings to test humanity. When Iolanthe was a teenager, the Jesuit Bertie de Montreuil was assigned to be her mentor, and while the young princess behaved with a similar willfulness and insolence to other royals Bertie had met, he found that she was both brilliant and talented. Through Bertie's teachings, Iolanthe came to learn more about the trials than most other royals, and was even privileged enough to see some secret artefacts such as the Vatican globe albeit only a brief glimpse of one side.
Iolanthe quickly became Bertie's best student, coming to see him as a surrogate father figure, and was given the nickname Io after Jupiter's fourth moon by him.
Throughout her years working at the Hall of Royal Records at St. Michael's Mount , Iolanthe was able to further learn more of the history and information that the Four Kingdoms had accumulated over the millennia, often reading or discovered documents containing new information that surprised her, such as a diary of Elizabeth I's mentioning a previously unknown chess championship in Constantinople.
Among the other things Iolanthe learned was the royals' records on the Machine and how its components were affected over the years, a letter to Queen Elizabeth from Sir Francis Drake noting that he had the coordinates to the Three Secret Cities of the Trial of the Cities , and a lost Kings Eyes Only document revealing that the Altar of the Cosmos had inadvertently been uncovered in As a high-ranking member of the British royals, Iolanthe was also allowed to access her family's talisman, one of the six Pillars - the Fourth - of the Machine.